EDUCATION
1997 M.D. Harvard Medical School
1995 Ph.D. Biochemistry, University of California, San Francisco
1988 B.A. Chemistry, Carleton College, Northfield, MN
RESEARCH
2006-Associate Professor of Cell Biology, Harvard Medical School
2000- Assistant Professor of Cell Biology, Harvard Medical School
1999- Head, Biological Technology, Institute of Chemistry and Cell Biology, Harvard Medical School
1997-2000 Institute Fellow, Institute of Chemistry and Cell Biology
1995-1997 Completion of medical studies, Harvard Medical School
1988-1995 M.D./Ph.D. Candidate, University of California, San Francisco. Dissertation Research with Marc W. Kirschner "Biochemical Characterization of the Cyclin B Ubiquitin Conjugation System"
1984-1988 B.A. Candidate, Carleton College. Undergraduate Reseach with Jerry Mohrig, Chemistry Department. "Stereochemistry of Enolate Protonation"
HONORS
2005 Harvard Medical School Prize for Excellence in Teaching (Years 1&2)
2005 Hellman Family Foundation Award
2004 Lynch Foundation Award
2001 Damon Runyon Scholar Award
2000 Harvard Armenise Foundation Award
1997 Institute Fellow, Institute of Chemistry and Cell Biology, Harvard Medical School
1992 Distinction in Teaching, UCSF
1988-1995 Medical Scientist Training Program, UCSF
1988 Honors in Independent Study, Carleton College
1988 Distinction in Major, Carleton College
1988 Summa Cum Laude, Carleton College
1988 American Institute of Chemists Student Research and Recognition Foundation Award
1987 Reeve Prize, Carleton College
1984 William Carleton Scholar